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Ground squirrels experience distinct events that divide each year into a long hibernation season, defined by when animals remain sequestered in their hibernacula, and a conversely short active season, when animals are active, above-ground on a daily basis. During hibernation, ground squirrels are heterothermic, altering their metabolism and body temperature between long periods two to three weeks of torpor and Free sex dating in anchorage ak 99508 arousals, when euthermic levels of body temperature briefly less than 1 day resume. However, male but not female ground squirrels return to continuous euthermy in spring for two to three weeks before first emerging to the surface, since sustained high body temperatures are necessary for gonadal growth and maturation [ 2728 ].

In contrast, females end heterothermy three to four weeks after males and begin euthermia and emerge within 3—4 days. Females re-enter hibernation in August, up to eight weeks before males [ 2526 ]; this results in active and hibernation seasons that can differ in length between the sexes by three to four months. There can be relative differences in annual timing associated with differences in habitat between populations as well. Using long-term data comparing free-living female AGS in two nearby populations in northern Alaska, we have shown that seasonal differences in snow-cover are associated with significant differences in their phenology [ 1629 ]. At one site, Atigun, where snowmelt occurs on average 26 days earlier than the other site, Toolik, female AGS emerge from hibernation, give birth and enter hibernation significantly earlier than females living at Toolik, only 20 km away [ 16 ].

Snow-cover in both autumn and spring may be major factors influencing female phenology [ 29 ]. To understand better the impact future changing patterns of snow-cover may have on AGS as a species, we investigated the long-term 13 years phenology of male AGS living at each site during similar time periods as our investigations of females. We compared phenological differences between the sites in relation to the calendar year, the timing of snow-cover, and in comparison with the emergence date of females.

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Atigun is slightly sloping along the northern bank of the Dzting River. The flora is dominated by a mix of low growing Salix sp. Thaw depth of the sandy soil reaches 1—2 m in August, into which AGS have dug a high density of burrows and nest sites and hibernacula. The Toolik area is categorized as cotton grass Eriophorum tussock and dry tundra with more than 98 per cent of the vegetative biomass and productivity attributable to only 10 species [ 33 ]. The topography is relatively flat, with gently rolling hills underlain by rocky soils with a depth of thaw usually less than 1 m [ 34 ]. AGS burrows are less dense and more isolated from each other at Toolik than at Atigun.

Entrances of AGS burrows at both sites are randomly oriented and showed no differences in exposure to prevailing environmental conditions, such as solar radiation, shade or wind. Owing to less overwinter accumulation of snow and high winds that disperse snow, spring snowmelt occurs on average 26 days earlier at Atigun than at Toolik, and in autumn snow-cover occurs on average 14 days later at Atigun than at Toolik. In men, testosterone is produced primarily in the testicles. Small amounts of testosterone are produced by the ovaries in women and the adrenal glands in both sexes. The portion of testosterone which is not bound is known as Free Testosterone.

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